Loc Start
Decrease font size  Default font size  Increase font size 


In press:

  • Holwerda, F., Beatty, B.L., Schulp, S. in press 2013. Dental macro- and microwear in Carinodens belgicus, a small mosasaur from the type Maastrichtian. Netherlands Journal of Geosciences.
  • Ribeiro, V., Mateus, O., Holwerda, F., Araujo, R., Castanhinha,R. in press 2013. Two new theropod egg sites from the Late Jurassic Lourinha Formation, Portugal. Historical Biology.

Presentations at meetings:


  • Holwerda, F., Beatty, B.L., Schulp, S. Dental microwear in the Maastrichtian mosasaur Carinodens belgicus. Poster presented at 72nd Society of Vertebrate Paleontology annual meeting, Raleigh, NC.
  • Ribeiro, V., Holwerda, F.  A new theropod eggsite in the Late Jurassic of Casal da Rola, Lourinhã, Portugal and the nesting preferences of basal theropod dinosaurs. Poster presented at 8th Encontro Nacional de Biologia Evolutiva, Oeiras, Portugal.


  • Holwerda, F., Pol, D., Gröcke D., Rauhut, R. Tooth morphotypes of sauropod dinosaurs from the Cañadon Asfalto Formation (Middle Jurassic) of Patagonia. Poster presented at 71nd Society of Vertebrate Paleontology annual meeting, Las Vegas, NV.


2013 PhD student in DFG project: Revision of basal sauropods from the Middle Jurassic of Patagonia and the early evolution of eusauropods. Supervisors: dr. Oliver Rauhut and dr. Diego Pol.
2012-2013 Universidade Nova de Lisboa, FCT research fellow in Project Dinoeggs: Dinosaur eggs and embryos from Portugal: paleobiological and paleoenvironmental implications. Supervisor: prof. dr. Octávio Mateus.
2011 Research Assistant at Durham University, Faculty of Earth Sciences, SIBL, Ocean Anoxic Events. Supervisor: dr. Darren Gröcke.
2010 MSc project Utrecht University and MEF Trelew, Argentina: ‘Middle Jurassic basal sauropod teeth from Patagonia, Argentina’. Supervisors: dr. Diego Pol and dr. Oliver Rauhut.